COLUMBUS, Ohio — The world’s first gorilla born in a zoo, a female named Colo who became the oldest known living gorilla in the United States, has died at age 60, the Columbus Zoo and Aquarium said Tuesday.

Colo died in her sleep overnight, less than a month after her birthday, the zoo announced. She surpassed the usual life expectancy of captive gorillas by two decades.

Colo had been on exhibit in view of visitors on Monday, when the zoo offered free admission for Martin Luther King Jr. Day. Fond of baseball caps, the gorilla was seen in her enclosure toting around a yellow cap with a gorilla on it, given to her by the zoo team caring for her.

The zoo said the gorilla’s body will be cremated and the ashes will be buried at the zoo.

Colo was born at the zoo on Dec. 22, 1956. She was a mother of three, grandmother of 16, great-grandmother of 12 and great-great-grandmother of three.

Associated Press